Moguera's name is derived from "mogura" meaning "mole", since it is a robot that is not only used for combat, but for digging purposes as well.
Moguera was a war machine created by the Mysterians. They sent Moguera to attack a village and destroy its shrine. Moguera moved up the river and was attacked by policemen. Moguera then destroyed a power plant, and then attacked a town. It was attacked by the military, but their weapons had no effect. Moguera was defeated when it tried to cross a bridge that was rigged to explode. Another Moguera was released and it attacked from underground. It dug up too close to a Markalite Cannon, and it fell on it.Godzilla Island, and was known as Proto-Moguera (プロトモゲラ?, Puroto Mogera) so that it wouldn't be mistaken for the other Moguera that appeared in Godzilla vs. SpaceGodzilla.
Appearing on the first world, the giant mech Moguera will follow Godzilla and Mothra through out the game, attacking them on each world. Moguera lacks versatility as he is only able to walk across the stage rather slowly, usually in response to the player's movement; in fact, if the player advances on Moguera there is a chance he will back himself into the corner. Moguera can jump, not very well however, but it will still be enough to knock Mothra to the ground if she is too close. While Moguera lacks the maneuverability of Gezora, he makes up for it with fire power as his Eye Beam will deal noticeable damage to either Godzilla or Mothra; furthermore, the beam is incredibly accurate and its very rare that either Godzilla or Mothra will be able to dodge it. Moguera has two variations on his Eye Beam, one a bright red and the other a brown; which he uses depends on who he is fighting against, bright red against Godzilla and brown against Mothra. The difference in the beams, however, is purely cosmetic.
Like the Heisei Moguera, the original Moguera did not have an actual roar, although its weapons and attacks did make sounds.
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