The Meganulons' name comes from Meganeura, a species of extinct dragonflies from the Carboniferous period approximately 300 million years ago.
Presumably, the Meganulons, like Rodan, are from the Cretaceous Period. They were awakened by mining operations in Kitamatsu village, Kumamoto Prefecture.
HistoryRodan. They were first awakened during a mining operation, and attacked and ate a few trapped miners. They escaped the mine and attacked a nearby village. A team was sent into the mine to exterminate the Meganulons. Shigeru, a member of the team, found an egg in the cave which hatched into Rodan. Rodan ate the Meganulons in the cave until it had enough strength to fly away. Tokyo when it started to ooze. The egg hatched into several Meganulons, which flooded the city and attacked and killed several citizens before molting into Meganula.
MeganulaThe Meganula (メガニューラ?, Meganyūra) were the second form of the Meganulon. They attacked Godzilla to steal his energy for their queen. Godzilla managed to kill most of the Meganula, but some escaped with his energy. They flew to Tokyo and gave the energy to Megaguirus, dying in the process.
In Rulers of Earth #17, Lucy Casprell and her team travel to Brazil, where a large number of Meganulon eggs have been uncovered in a nearby rainforest. As Lucy and her team investigate the eggs, a claw emerges from one of the eggs.
In Godzilla: Cataclysm #3, the Meganula arrive in the refugees' village along with Megaguirus in order to feed on Biollante, who is still regenerating from her battle with Godzilla. Mothra arrives to protect Biollante and battles Megaguirus. The Meganula attempt to swarm and bite Mothra, but are knocked away by her wings.
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